Characterizing dog cognitive aging using spontaneous problem-solving measures: development of a battery of tests from the Dog Aging Project

We thank Canine Companions and Sit! Stay! Play! for facilitating and supporting research at their facilities. We thank Chloe Burkholder, Elizabeth Carranza, Grace Davis, Emily Davis, Abby Flyer, Catherine Fry, Gabi Garcia, Ana Gonzales, Emily Hilyard, Emma Kristal, Kevin McLaughlin, Belinda Morton, Tyler Rodriquez, Alexa Sell, Caitlyn Seymour, Olivea Spencer, Brianna Tamayo, Natya Thirunagari, Christina Cawood, Alessandra Ostheimer, Angelina Schirle, Rachel Shaberman, Olivia Gittings, and Nova Vogel for their help with research logistics, data collection, and/or video scoring. Lastly, we thank the many individuals and dogs in our community who participated in these studies.

Dog Aging Project consortium authors Joshua M. Akey1, Brooke Benton2, Elhanan Borenstein3,4,5, Marta G. Castelhano6,7, Amanda E. Coleman8, Kate E. Creevy9, Kyle Crowder10,11, Matthew D. Dunbar11, Virginia R. Fajt12, Annette L. Fitzpatrick13,14,15, Unity Jeffery16, Erica C. Jonlin2,17, Matt Kaeberlein2, Elinor K. Karlsson18,19, Kathleen F. Kerr20, Jonathan M. Levine9, Jing Ma21, Robyn L. McClelland20, Daniel E.L. Promislow2,22,23, Audrey Ruple24, Stephen M. Schwartz14,25, Sandi Shrager26, Noah Snyder-Mackler27,28,29, M. Katherine Tolbert9, Silvan R. Urfer2, Benjamin S. Wilfond30,31

1Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, NJ, USA.

2Department of Laboratory Medicine and Pathology, University of Washington School of Medicine, Seattle, WA, USA.

3Department of Clinical Microbiology and Immunology, Faculty of Medical and Health Sciences, Tel Aviv University, Tel Aviv, Israel.

4Blavatnik School of Computer Science, Tel Aviv University, Tel Aviv, Israel.

5Santa Fe Institute, Tel Aviv University, Santa Fe, NM, USA.

6Cornell Veterinary Biobank, College of Veterinary Medicine, Cornell University, Ithaca, NY, USA.

7Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, NY, USA.

8Department of Small Animal Medicine and Surgery, College of Veterinary Medicine, University of Georgia, Athens, GA, USA.

9Department of Small Animal Clinical Sciences, Texas A&M University College of Veterinary Medicine & Biomedical Sciences, College Station, TX, USA.

10Department of Sociology, University of Washington, Seattle, WA, USA.

11Center for Studies in Demography and Ecology, University of Washington, Seattle, WA, USA.

12Department of Veterinary Physiology and Pharmacology, Texas A&M University College of Veterinary Medicine & Biomedical Sciences, College Station, TX, USA.

13Department of Family Medicine, University of Washington, Seattle, WA, USA.

14Department of Epidemiology, University of Washington, Seattle, WA, USA.

15Department of Global Health, University of Washington, Seattle, WA, USA.

16Department of Veterinary Pathobiology, Texas A&M University College of Veterinary Medicine & Biomedical Sciences, College Station, TX, USA.

17Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, WA, USA.

18Bioinformatics and Integrative Biology, University of Massachusetts Chan Medical School, Worcester, MA, USA.

19Broad Institute of MIT and Harvard, Cambridge, MA, USA.

20Department of Biostatistics, University of Washington, Seattle, WA, USA.

21Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, WA, USA.

22Jean Mayer USDA Human Nutrition Research Center on Aging, Tufts University, Boston, MA, USA.

23Department of Biology, University of Washington, Seattle, WA, USA.

24Department of Population Health Sciences, Virginia-Maryland College of Veterinary Medicine, Virginia Tech, Blacksburg, VA, USA.

25Epidemiology Program, Fred Hutchinson Cancer Research Center, Seattle, WA, USA.

26Collaborative Health Studies Coordinating Center, Department of Biostatistics, University of Washington, Seattle, WA, USA.

27School of Life Sciences, Arizona State University, Tempe, AZ, USA.

28Center for Evolution and Medicine, Arizona State University, Tempe, AZ, USA.

29School for Human Evolution and Social Change, Arizona State University, Tempe, AZ, USA.

30Treuman Katz Center for Pediatric Bioethics, Seattle Children's Research Institute, Seattle, WA, USA.

31Department of Pediatrics, Division of Bioethics and Palliative Care, University of Washington School of Medicine, Seattle, WA, USA.

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