We thank the staff of the imaging core at First Core Labs, National Taiwan University for technical support as well as members of Liu and Chang laboratories for helpful comments.

This work was supported by Ministry of Science and Technology grant MOST 107-3017-F-002-002 to Y.W. Liu, MOST 110-2636-B-007-011 and 111-2636-B-007-009 to Y.C. Lin, and National Taiwan University grant NTU-CDP-112L7809 to Y.W. Liu. J.M. Baskin acknowledges support from a Beckman Young Investigator award, a Sloan Research Fellowship, and the National Science Foundation (CAREER CHE-1749919). R. Tei was supported by Funai Overseas and Cornell Fellowships. Open Access funding provided by the National University of Taiwan.

Author contributions: Conceptualization: Y.-A. Su, H.-Y. Chiu, Y.-C. Chang, Y.-W. Liu, and Z.-F. Chang; Methodology: Y.-A. Su, H.-Y. Chiu, Y.-C. Chang, Y.-C. Lin, R. Tei, H. Bauer, Y. Feng, and J.M. Baskin; Investigation: Y.-A. Su, H.-Y. Chiu, Y.-C. Chang, C.-W. Chen, S.-S. Lin, X.-R. Huang, H.-C. Wang, C.-W. Chen, and J.-C. Hsu; Supervision: Y.-C. Lin, J.M. Baskin, Y.-W. Liu, and Z.-F. Chang; Writing—Y.-C. Chang, H.-Y. Chiu, and Y.-A. Su; Writing—review & editing: Y.-A. Su, Y.-W. Liu, and Z.-F. Chang.