Mammalian Behaviour and Evolution Group, Institute of Infection, Veterinary and Ecological Sciences, University of Liverpool, Leahurst Campus, Neston CH64 7TE, United Kingdom

*Corresponding author. Present addresses: Konrad Lorenz Institute of Ethology, Department of Interdisciplinary Life Sciences, University of Veterinary Medicine Vienna, Savoyenstrasse 1, 1160 Vienna, Austria; Department of Behavioral and Cognitive Biology, University of Vienna, University Biology Building (UBB), Djerassiplatz 1, 1030 Vienna, Austria; email: [email protected].

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